Table Of ContentShizuoka,Mar21,I992
InMemoriam Keiichi KusAMA
(6Feb.1924-6cot.1998)
Apageof thehistoryof theJapanesecoleoptero1ogywasclosedonOctober6,
1998,whenwelostKeiichiKUsAMA,D.Sc.,thefounderandthefirstpresidentofthe
JapaneseSocietyofColeoptero1ogy.Hesuffered fromstomachcancer forsometjme,
butkeptthesecrettohimselfanddidnotmakeusknownabouthisseriouscondition.
Hewas74yearsoldatthattime.
KuSAMAwasbornatTokyoonFebruary6,1924,butwassoonmovedtoZushj,a
WellknownresorttownonthePacific,mainlyforhishealth.Therehedevelopedinter_
estincollectinginsects,andaboveallincerambycidbeetlesafterthefifthyearofthe
ShigeoTsUYUK1andMasatoshiTAKAKUwA
elementaryschool. Thiscontinuedthroughhisstudent daysatShonanHighSchool
andtheFacultyofScience,UniversityofTokyo,thoughhestudiedbiochemistryinthe
latter. Hefinishedthepostgraduatecoursein1954, and receivedhisSc.D. in 1960
fromthesameuniversity.In1963,hewasinvitedfromShizuokaUniversityasanasso-
ciateprofessorattheDepartmentofHumanitiesandSciences,andwaspromotedtoa
professorofsciencein1967.Afterthat,helivedall thetimeinShizuokaCity,even
afterhisretirement in1988.
Thus,KusAMAwasnotaprofessionalentomologistbutmainlystudiedradioactive
isotopesintheuniversity.However,hewrotemanyacademicandnon-academicpapers
ontheCerambycidae.Alreadyinhisstudentdays,hecompiledseveral listsoflocal
cerambycid faunas. In1970,hefirstpublishedadescriptionofanewspecies, Necy-
dalismoriyat fromAmami-0shimaIslandofthecentralRyukyus,sincenecydalinece-
rambycidswereoneofhisfavourites.HecompiledlistsoftheJapaneseCerambycidae
threetimes,in1959,1965and1973,andgaveevaluationbysixrankstoallthenative
speciesasregardsoccurrence frequencyof theiradults:verycommon,common,not
common, few,rare,andveryrare.Theselistswereveryuseful forbeginnersnotonly
togiveinformationonecologicalanddistributionaldataofJapanesecerambycids,but
toarousetheirinterestinstudyingthem.Astheresult,thecerambycidfaunaofJapan
wasrapidly clarified inevery respect - taxonomy, phylogeny, distributions,habits,
hostplants,andsoon.
KusAMA'scontributionstotheJapanesecoleoptero1ogycanbedividedintothree
categories: taxonomicalstudies,establishmentofasocietydevotedtothecoleoptero1-
ogy,andpromotionofamateur1ongicornists.Hispaperspublishedbeforehisretire-
mentwerealreadyenumeratedonpages3-6inavolumeentitled“Kusamaia”,which
waspublished in 1988 as No 3oftheSpecialBulletinof theJapaneseSocietyof
Coleoptero1ogy,sothatthereisnoneedtorepeat.However,specialmentionshouldbe
madeofthebook“TheLongicorn-BeetlesofJapaninColor'(1984).Thiswasthefirst
reviewofalmostallthespeciesoftheCerambycidaeoccurringinJapan,andthevol-
ume numbering 566 pages with many colour illustrations was completed under
KUsAMA'ssupervision.
KUsAMAactivelyjoined intheestablishmentofour society for developingthe
coleoptero1ogyinJapan,andservedasthefirstpresidentfrom1973tot987.Underhis
presidency, thesocietymadeasteadydevelopment,andaftertheamalgamationwith
theColeopterists'AssociationofJapanin1989,hasattainedthepresentprosperity.He
wasanhonorarymemberofthesocietyinhislastyears,butwasstillactiveinevery
way.
KUsAMAwasalwayskindtoyoungcoleopterists,particularlytothosewhowished
tostudy cerambycidbeetles. Underhiscompetentguidance, they havegrownup to
fledged1ongicornists,someofwhomgainleadingpositionsinthefieldofcoleoptero1-
ogyinJapan.Healwayswishedtoutilizehiscollectionofspecimensandliteraturefor
theprogressof thecerambycido1ogy.Complyinghiswishes,wedecidedtokeep,at
least forthetimebeing,hisspecimensoftheCerambycidae,mainlycollectedinJapan,
In Memoriam Keiichi KusAMA
Southeast Asia, North AmericaandNew Guinea, at thehandsofShigeo TsUYUKl,
ZushiCity,andliteratureofentomologyatthehandsofMasatoshiTAKAKUwA,Kana-
gawaPrefecturalMuseumofNaturalHistory,0dawara,bothKUsAMAs' studentsinthe
cerambycido1ogy. Wesincerely hope that these inheritances will proveuseful for
youngergenerations.
Wewarmlydedicatethisfirst issueofthe1999Elytra tothelateProfessor Dr.
Keiichi KUsAMA.Fourteenpapersdedicatedtohismemorywillappearonthefollow-
ingpages.Theyarewrittenbyhisstudentsinthecoleoptero1ogyandhissuccessorsin
thepresidencyofthesociety.
ShigeoTsUYUKI
FormerVice-president
and
Masatoshi TAKAKUwA
PresentVice-president
Elytla,「o;kvo,27(1):5-10,May15,l999
TwoNewSpeciesofCerambycidBeetlesRelatedtoE:x:ocentrusfisherz
(Coleoptera,Cerambycidae,Lamiinae) fromTaiwan
ShigeoTsUYUKI
7-1-24,Zushi,Zushi-shi,KanagawaProf.,249-0006Japan
Abstract Twonewspeciesof thecerambycidgenusE?ocentrusaredescribed
fromTaiwanunderthenamesE kusamaispnovandEke11chiisp nov.,bothofwhichre-
semble in coloration E fis/lei・tGREsslTT, 1935 fromFar Eastern Asia. The former
markedlydiffersfromE. fshe,・iinthemalegenitalicfeatures,whilethelatterisveryclose
ite・,1.
te . fs
Thelamiinegenus Ex:ocentrus of Taiwanwas first reviewedby KUsAMA and
TAHIRA(1978),and13speciesincluding7newtaxawerelistedthere.After then,only
onespecieswasaddedtotheTaiwanesefaunabyMAKIHARA(1986) fromLanyuIsland
offsoutheastern Taiwan. However,thelateProf. Dr. Keiichi KusAMAseems tohave
foundoutoccurrenceofstillmoreExocentrusspeciesinTaiwan,becausesomeunde-
termined specimens with red labels werepreserved inhisprivatecollection. I am
goingtodescribetwonewspeciesfromthemandtonameboththespeciesa量erhim.
Thetwoho1otypesdesignatedinthispaperwillbedepositedinthecollectionof
theNationalScienceMuseum(Nat. Hist),Tokyo.Theparatypesarein thelateProf.
Dr.KeiichiKUsAMAs' privatecollection.
Beforegoing further, I wish toexpress my sinceregratitude to Dr. Masatoshi
TAKAKuwAoftheKanagawaPrefecturalMuseumofNaturalHistory,0dawara, forhis
kindhelpinpreparingtheoriginalmanuscriptofthispaper.ThanksarealsoduetoMr.
ToshioKOBAYASHIofYokohamaforsupplyingmewithvaluablematerialsusedinthis
paper.
Thisshortpaperisdedicatedto thememoryofmylatemaster, KeiichiKUsAMA,
whoconstantly ledmetothe1ongicorn-beetleworldofgreat interest throughmore
than45yearssincethetimeofmysenior-highschoolstudent.
E;x:ocentruskusa''taisp nov.
(Figs.1,2,7-l2)
Male. Small, castaneoustodark castaneous; headalmostblackish, though the
clypeusand theanterior partof labrumareyellowishbrown; lateralprojectionsof
pronotum,humeri,apicalpartofelytraandclawsbrown.
Head finely and densely punctate, densely clothed with pale white recumbent
ShigeoTsUYUK1
2
Figs. 1-6. HabitusofExocen1t・usspp. - l,E kusa,naisp nov., holotype;2,same, ,paratype;3,
E kelichiisp nov., holotype;4,same,9,paratype;5,E.fisheriGREssrrT,(3, fromFukushimaPre-
,
fectureofcentralHonshu;6,samespecies, fromthesamelocality.
,
scale-likepubescence,andalsosparselywithvery longsemirecumbentbristles; frons
smoothly and distinctly swollen; eye large, fully prominent, coarsely faceted, with
lowerlobenearlycircularthoughslightlyhigher thanwide.Antennaexceedingelytra1
NewRelativesofE?-oce,ll,-tlsfis/1e,'i
I
8
Figs. 7-l2. E1ocent,・llsktlsanla1sp nov., holotype. - 7,Pronotum;8, 8thabdominalsternite in
innerview;9,medianlobeinlateralview;10,apexofthesame,ventralview;11, tegmeninventral
view;12,same,lateralview
apexatapical1/2of8thtobaseof9thsegment;scapeweaklyclavate,gentlyswollen
tobehind themiddle,sparselywitherect orsemierect longbristles;segments2-10
withsparselongbristlesbeneath;relativelengthsofsegmentsintheholotypeas fol-
lows: - 4.4: l: 3.9: 3.8: 3.2: 3.1: 2.9: 2.8: 2.6: 2.4: 2.4. Pronotum fairly transverse,
about148timesaswideaslengthexcludinglateralprojections;basedistinctlyshorter
thanapex;dorsumpunctateandhairednearlyasonhead; lateralprojectionsslender
andstrongly oblique.Scutellumwider thanlong(1.7:1),subtriangularwith rounded
apex.Elytra2.13timesaslongashumeralwidth,almost twiceaslongasmaximum
width,stronglyelevateddorsad,highestbehindthemiddle;sidessinuate,convergent
frombehindhumeritobasal3/10wheretheyareweaklyconstricted,gentlydivergent
fromtheretobasal2/3whichisthewidest,andthenarcuatelyconvergentapicadin
dorsalview;eachapexnarrowlytruncate;surfaceratherdenselyanddeeplypunctate,
denselyclothedwithwhitishsemirecumbentscales,andalso relativelydenselywith
erectorsemierectstout bristlesallover;humeral brownpartmorevivid thaninapex.
Abdomen densely clothed with recumbent palewhitish pubescence; anal segment
somewhatdenselywithlongsemirecumbentbristlesonaboutapicalhalf,broadlyand
shallowlyemarginateatapex;8thsternitehemie11iptica1,withafewshorthairsatapex.
Hindfemoraclavate,eachgraduallyswollenfrombehindthemiddle.
Genitaliastout;medianlobethick,stronglybentventrad,withdorsalplateappar-
ently shorter than theventral; tegmen broa smoothlycurved in lateral view, with
parameresshortandseparated.
Female. Antennaexceedingelytralapex at9thsegment.Elytrabroader,2.08
timesaslongashumeralwidth,1.92timesaslongasmaximumwidth;sidesalmost
straightlydivergent frombehindhumeritobasal3/5,thenarcuatelyconvergentapicad
indorsalview;apexweaklyrounded.Analsegmentelevated;apextriangularly,deeply
emarginateatthemiddle.Hindfemoraalmostsimilartothoseofmaleinshape.
ShigeoTsUYUK
Length:(:S5.2-5.5mm; :6.0-6.3mm.
Typeseries. Holotype: dSankuang,TaoyuanHsien,centralTaiwan,1-V-1979,
S.TSUYUKIleg.Paratypes:2(5d3 ,samelocalityandcollectorastheholotype.
Distribution. Central Taiwan.
Atfirstsight,thisnewspeciesresemblesE.fisheriGREsslTT,1935fromFarEast-
ernAsiainthecoloration,butevidentlydiffersfromitinthefeaturesofpronotum,the
8thabdominalsterniteandgenitaliaasdescribedabove.
Exocentruskeiichiispnov.
(Figs3,4,13-19)
Male. Relatively large,darkcastaneoustoblackish;clypeus,anteriorpart of
labrum,mouth-partsexcept formandibles,andgulabrown;humeri,andlateraland
apicalpartsofelytrabrowntoyellowishbrown;mandibles,spursandclawsreddish
brown
Headfinely,denselypunctate,denselyclothedwithpalerecumbentpubescence,
andalsosparselywithverylongsemirecumbentbristles; fronsslightlyswollenbilater-
ally;eyelarge, fullyprominent,coarselyfacete withlowerlobecircularwithstraight
innermarginthoughhigherthanwide.Antennaexceedingelytra1apexatapicalpartof
8thsegment;scapeweaklyclavate,gentlyswollentobehindthemiddle,sparselywith
longinclinedbristles;segments2-10withratherdenseinclinedbristlesbeneath; rela-
tivelengthsofsegmentsin theholotypeas follows: - 5.0: 1: 5.6: 5.3:4.7: 4.6: 4.l:
3.9:3.7:3.4:3.6.Pronotum fairlytransverse,about143timesaswideaslengthex_
I
一
( 16
一
14 一 ーー 一- 15
FigS. 13-l9. Exoce'It''uskeilchiisp nov.,l3,holotype. - l3,Pronotum;14,7thabdominal tergitein
innerview; l5,8thabdominalsterniteininnerview;16,apicalpartofmedianlobeinlateralview;17,
same,ventralview;18,tegmeninventralview;19,same,lateralview.
NewRelativesofExocen1t・us/ls/1e・,t
eluding lateralprojections; base longer thanapex (1.2:1); lateral projectionsthick
withroundedapexandmoderatelyoblique;dorsumpunctateandhairednearlyason
headthoughthebristlesareshorterandpalewhitishscale-likehairsarepresentonthe
medianlongitudinalzoneandnearthebase.Scutellumwiderthanlong(1.48:1),para_
helical.Elytra2.10timesaslongashumeralwidth,justtwiceaslongasmaximum
width,weaklyelevateddorsa highestbehindthemiddle;sidesverygraduallydiver-
gent tothemiddlewhich isthewidest, thenarcuatelyconvergent apicad indorsal
view;apicesroundedor narrowlytruncate;surface finelypunctate, denselyclothed
withwhitishpalepubescenceandalsorelativelydenselywitherectorsemirecumbent
stoutbristlesallover;humeralbrownpartmorevividthaninlateralandapicalparts.
Abdomendensely clothedwithrecumbent palepubescence, andalsosparsely with
longhairsalmostallover;analsegmentsomewhatdenselywithlongsemirecumbent
hairsonaboutapicalhalf,broadlyandshallowlyemarginateatapex;7thtergitetrans-
verselyparabolical,withslightlyemarginateapex;8thsterniteshapedasaratherbroad
isoscelestriangle.Hindfemorawellclavate,eachabruptlyswolleninapical2/3.
Genitaliaslender;medianlobeweaklybentventrad,withdorsalplateapparently
shorter than theventral; tegmensigmoidallycurvedjustbeforethemiddleinlateral
view,withparameresstoutandseparated.
Female. Antennaexceedingelytra1apexat9thsegment.Elytra2.19timesas
longashumeralwidth,1.94timesaslongasmaximumwidth;sidesalmoststraightly
divergent frombehindhumeri tobasal t/2, thengraduallyandarcuatelyconvergent
apicadindorsalview.Analsegmentelevated;apexsinuatelyemarginateatthemiddle.
Hindfemoraweaklyclavate,eachgraduallyswollenfrombasetobehindthemiddle.
Length: d5.5-6.8mm;e:64-7.0mm.
I
-
一
23
一
21 22
Figs.20-26. litocent''fisfs1/e,・iGREssITT, from Fukushima Prefecture ofcentral Honshu. - 20,
,
pronotum;21,7thabdominaltergiteininnerview;22,8thabdominalsterniteininnerview;23,apical
partofmedian1obeinlateralview;24,same,ventralview;25, tegmeninventralview;26,same,lat-
eral view.
10 ShigeoTsUYUK1
Typeseries. Holotype: Sankuang,TaoyuanHsien,centralTaiwan,4~7-V-
,
I978,T.KOBAYASHIleg.Paratypes:5(;(3,6 ,samedataastheholotype.
fs lbuf1on. Central Taiwan.
ThepresentnewspeciesisverycloselyalliedtoE ftsheriGRESSITT, l935from
FarEasternAsia,anddoubtlesshasaverycloserelationshipwithitamongthehitherto
knownmembersofthegenus,becauseofthemorphologicalsimilaritynotonlyofex-
ternalcharactersbutalsoofgenitalicfeatures.However, theTaiwanesespeciesdefi-
nitelydiffersfromtheFarEasternAsianoneinthe followingpoints:1)elytra1punc-
turesdistinctly finerandshallower,2)analsternitemoretransverse,withshallowly
emarginateandmoredenselyhairyapex,3)8thabdominalsterniteratherbroadisosce-
lestriangular,4)dorsalplateofmedianlobeofmalegenitaliadistinctlyshorterthan
theventral.
要 約
露木繁雄:ヨコグロケシカミキリに似た台湾産アラゲケシカミキリ属の2新種. - 故草
間慶一博士が研究中であった台湾産ヵミキリムシ科甲虫のうち, アラゲケシカミキリ属の2新
種, Ex:ocentruskusa,naispnov.とE kelichiispnov.を記載した. いずれも色彩的には, 極束地域
に分布が知られるョコグロケシカミキリに似てぃる. しかし, 前種はそれとは系統的にかなり
異なった種で, 前胸背板をはじめ鞘翅, 第8腹板, 交尾器の形状がまったく違つてぃる. 後
種は, 外部形態ばかりでなく,SI交尾器の構造も, ヨコグロケシカミキリと基本的に同じであり,
きわめて近縁のものであることには疑いの余地がないが, 鞘翅の点刻の状態や腹部末端節, 第
8腹板の形状, さらに 交尾器中葉片の背板が腹板より明らかに短い点などが異なるので, 別
種として扱った.
References
GREsslTT,J. L., l935. NewJapanese1ongicornbeetles(Col:Cerambycidae). Kontyu, 「okyo,9:166-
179.
- 1951. LongicornbeetlesofChina. Longicornia,2: l-667,22pls.
KUsAMA,K.,&Y.TAHIRA,1978. ThegenusExocentrusMULsANTofJapananditsadjacent「egiOnS:(2)
therevisionofTaiwanesespecies. Elytra,'nokyo,6:9-32.
MAK田ARA,H., & T.NllsAT0, 1986. Some10ngicornbeetlesfromTaiwan(C0leoptera,Ce「ambyCidae),
withdescriptionsofthreenewspecies. Misc.Rcpt.HiwaMusflat.Hist.,(24):1-17.
NAKAMURA,S.,H.MAKIHARA&A.SAIT0, l992. Check-listof1ongicorn-beetlesofTaiwan. MiSC.Rcpt.
HiwaMus nat.Hist.,(30):1-126.
