Table Of ContentNewsletter of the Colorado Native Plant Society
dedicated to the appreciation and conservation of the Colorado native flora”
.
Volume 24 Number 5 November—December 2000
Generic Changes to the Scrophulariaceae affect the Rocky Mountain Flora
Dr. Neil Snow, Biological Sciences, University of Northern Colorado
A considerable nutnber of generic transfers from the incorrectly imply some kind ofprogress.
Scrophulariaceae (figwort family) to the Orobanchaceae (broom- The second principle of phylogeneticsjs that only monophyletic
rape family) and Plantaginaceae (plantain family) are now well (mo-no-fy-LEH-tik) groups should be giventaxonomic names. A
supported by new research. Herein, I provide an overview ofthe monophyleticgroupisonethatincludesanancestraltaxonandall
current philosophy underlying classification and the advent of (not just some) of its descendant taxa. In the past, taxonomic
molecular sources of data in systematics, both of which are sig- names have been given to groups thatare notmonophyletic. One
nificantly affecting the taxonomy of one of Colorado’s more example is Class Reptilia. Zoologists historically excluded birds
'irominentplantfamilies. fromReptilia,whichinsteadwere placedintheirownClassAves,
^'^nJlassification. The goal ofmodem taxonomy is to group organ- However, sincebirdsarealsodescendedfromthecommonances-
ismsinsuchawaythattheclassificationreflectsevolutionaryhis- torofotherreptiles,itis incorrectunderphylogenetic systematics
tory. The prevailing taxonomic school of thought among profes- to recognize Reptilia, without also including the birds. Stated
sional taxonomists is “Phylogenetic Systematics,” or simply anotherway.ClassReptiliabecomesmonophyleticifthebirdsare
—
“Phylogenetics.”Thisphilosophy was firstdeveloped indetail by included this is the preferred phylogeneticclassification.
the late German entomologist Willi Hennig. Although the initial ... systematics is a science and must reflect
embrace of phylogenetics was slow and considerable debate
ensued over its relative merits, phylogenetic systematics is now well-supported data
firmlyestablishedasthemostscientificapproachtoclassification.
Phylogenetic classification has twoover-riding principles. Moleculardata. Soonaftertheriseofphylogeneticsystematics,use
The first is that taxa are grouped together only by evolutionarily ofmoleculardataexp^ded rapidly. CertaincharacteristicsofDNA
derived characters (traits) that are sharedby twoormore species. found inthenucleus, mitochondria, andchloroplastscanbeusedas
Forexample, the presence offlowerscannot be used to helpclas- taxonomicmarkerstoassistintheprocessofclassification,although
sifymembersofthegenusCastillejawithinthepaintbrushes.This mitochondrial DNAis usedprimarilyforanimal classifications.
isbecauseall angiospermshaveflowers,andflowersthereforeare Moleculardata have enabled some previous taxonomic questions
not a derived character among members of Castilleja. However, to be answered satisfactorily. One example bears on relationships
the presence offlowers is aderived characterrelativeto all ferns, tothe grassfamily.Anumberofcandidatefamilies had beenpro-
whichreproduceonlyby spores. Inthis simpleexample,then,the posed as being mostclosely related tothe Poaceae. But none had
presence of flowers could be used to group together species of “Scrophulariaceae”continues on page 2
Castilleja relative to ferns, since flowers are evolutionarily
Contents
derived relativeto spores. Theword systematists usefora shared
derived character is “synapomorphy” (pronounced sin-AP-oh- About the Society .7
morf-ee). In contrast, an ancestral trait is a “sympleisiomorphy” Announcements ..... ....... .2, 3, 4, 6
(sim-PLEASE-ee-oh-morf-ee). These terms should be learned by Articles .1-2, 5
hose interested inthe moretechnical aspects of taxonomy, since
Calendar .8
^ney are used regularly by taxonomists. In contrast, “advanced”
(for derived) and “primitive” (for ancestral) are falling out of Chapter News and Announcements ... .2, 6
favor, particularly since “advanced” has a historical tendency to Society News and Announcements .... .3-4
Page 2 Aquilegia Vol. 24 No. 5
“Scrophulariaceae*"continued frompage 1 existing floras recognize these groupings, future publications will
been shown convincingly as more closely related than others recognize many ofthe newer concepts. Old habits, however, can
using traditional sources of data, such as morphology, anatomy, be hard tobreak. -
andchromosomaldata.Amolecularstudypublished several years Anumberofyears ago we tried toconvince my GrandmaAuerin^^
ago demonstrated convincingly that the Joinvilleaceae, which upstate New York to try a peanut butter andjelly sandwich. She
consists ofonlyoneortwospecies widely scattered inthe Pacific stated that humans are creatures of habit, and a lifelong habit of
region, is the sister family to the species-rich and ecologically hers was to politely decline peanut butter andjelly when offered.
dominant grasses. Associating certain generawithparticularfamilies is alsohabitual
Changes to the Scrophulariaceae. for botanists. Some may therefore resist the generic reconfigura-
Historically, the Scrophulariaceae has tionsoftheScrophulariaceae,Orobanchaceae,andPlantaginaceae.
been recognized as consisting of However, systematics is a science and must reflect well-support-
some 220 genera, which are particu- ed data. Amore accurate understanding ofphylogenetic relation-
larly common in temperate and tropi- shipshelpsus betterunderstandourfloristicheritage. Namesmat-
cal mountains. Genera such as ter in some ways, but not in others. After all, the monkeyflower
Castilleja, Mimulus, Pedicularis, adorningthemountain stream nearyourfeetduringalunchbreak
Veronica, and several others are still looks spilfy, irrespective of its family placement or whether
prominent in our area. Penstemon your sandwich contains peanut butterandJelly.
even has its center ofdiversity in the
mountainous areas of the western United States. However, the WESTERNSLOPEWORKSHOPS
Scrophulariaceae was notmonophyletic in its previous classifica- ANNOUNCED
tion. It probably comes as a surprise to many CoNPS members
GayAustin, PlateauChapter
that recentresearch has convincingly indicated the need to trans-
fermost generacomprisingthe Scrophulariaceaeelsewhere. For 17 seasons, CoNPS workshops have bfought native plant
In 1999, Drs. WalterJudd, ChristopherS, Campbell, ElizabethA. lovers togetherwithawell-informedinstructor. Theopportunity
Kellogg, and Peter F. Stevens published Plant Systematics: A to receive one-on-one instruction and informative lectures has
PhylogeneticApproach (SinauerAssociates, Inc.), which reveals made the workshop series one ofthe most popular Society pro-
thefollowing generic changestothe Scrophulariaceae. This book grams. This season, the Plateau Chapterofthe Colorado Native
is highlyrecommendedforthose interested inmore details about PlantSocietyis proudtoannounce several workshopstobecon^
phylogeneticsandtheimportanceofmoleculardata,aswell asthe ducted on the Western Slope. Attendees need only a love of
specific reasons for these changes to the Scrophulariaceae, My plants and a desire to learn about them. To registerfor Western
thanks to Dr. Richard Olmstead (University of Washington) and Slope workshops or for more information, please contact Gay
Dr. Judd (University of Florida) for sharing information on this Austin athome (970) 641-6264orwork(970) 642-4406.
recentclassification. How to Know the Families ofthe Western Slope
Analysis ofboth molecularand non-moleculardata suggests that Leaden Dr. RobinBingham
the following generafound in Coloradoformerly included in the Location: Delta
Scrophulariaceae should be placedasfollows; Date: January 27,2001
Scrophulariaceae: VerbascumandScrophularia. Inits newer Dr. Robin Bingham, Botanist at Western State Col|ege, has
and more restricted sense, the family now includes only the gearedthisworkshoptowardthebeginnerbotanistorthemember
genera containing non-sagittate anthers, in which the anther who would like to review! As such, this workshop will focus on
loculesareconfluentand openby a single distal slit. how to recognize the major plantfamilies ontheWesJein Slope.
Orobanchaceae: Castilleja, Pedicularis, Agalinis, Dr. Bingham will alsodiscuss basictermsforplantidentification
Orthocarpus, and Cordylanthus, The Orobanchaceae in its and how totell monocotsfromdicots. PleasecontactGayAustin
newer sense includes the hemiparasitic genera formerly athome(970)641-6264forworkshopregistration, whichis$12.
placedin the Scrophulariaceae.
Grasses ofthe Western Slope
Plantaginaceae: Antirrhinum, Besseya, Callitriche,
Leaden Dr. DavidL. Buckner
Ghionophila,Collinsia,Digitalis,Hippuris,Limosella,Linaria,
Location: DeltaorGrandJunction
Lindernia, Mimulus, Penstemon (including Leiostemon),
Date: March 17, 2001
Rhinanthus, Veronica (including Pocilla), and Veronicastrum.
The Plantaginaceae, thus, now includes the autotrophic Dr. Buckner, Consultant in Plant Ecology, Revegetation, and
“scrophs.” The aquatic genera Callitriche and Hippuris are Restoration with ESCOAssociates, will leada basic grass iden-
transfersfrom theCallitrichaceaeandHippuridaceae. tification class emphasizing grasses ofthe Western Slope. Tribe
characteristics, specifically, will be covered in this workshop,
Other changes in classification based on phylogenetic analysis
which will begearedtowardbothbeginnersandadvancedmem-
eventually will bemadeas morestudiesare published. Forexam-
bers. Microscopes and specimens will be available. Please con-
ple, somedata supportthemergingoftheAceraceae (maplefam-
tact GayAustin at home (970) 641-6264for workshop registra-
ily)intotheSapindaceae(soapbenyfamily).Althoughnoneofthe
tion, which is $12.
-
Vol. 24 No. 5 Aquilegia Page 3
NEWS AND ANNOUNCEMENTS
SOCIETY
Minutes of the CoNPS Board Meetings September 23, 2000
Alice Guthrie, CONPS Secretary New Business: Officers Jill Handwerk (President), Dave
Anderson (Vice President), Denise Culver (Treasurer), andAlice
September22, 2000 Guthrie (Secretary) were approved by the Board, with the next
Iteasnrer’s Report: Denise Culverdistributedthe budget report meeting scheduled for November 4, 2000 in Longmont Public
year-to-date, as well as the profitand loss statement. Denise also Safety Building, Conference Room, 225 Kimbark.
reviewed chapter subventions, calculated as 25 percent of mem-
bershipduesforeachchaptermember. Help!
Committee reports: Tllhairs Bob Clarke (Conservation), Sue TheBoardis requestingyourhelpwith thefollowing:
Martin (Membership), and Rick Brune (Field Trips) are stepping Donation of a folding table, 5-6 feet long, for Society dis-
done, with Velma Richards (Publications) cutting back on her plays. Please contactAlice Guthrie at (303) 651-3127.
responsibilities. Eric Lane will take over as Chair of Volunteeropportunities:
,
Membership, while JeffDawson will chairPublications, work- ProGreenShow inDenver,January2-5, 2001.Volunteers
ingwithVelma.TheChaifofPublicity is still vacantandthe sta- areneededtosetupandstafftheboothforeitherahalfday
tus of the Field Studies Committee is under discussion. Leo orwhole day. ContactJill Handwerkat (970) 491-5857.
BruederleoftheNewsletterCommitteereportedthattherewillbe Committee Chairs (andmepibers) are neededforthefol-
five regular issues ofAquilegia in 2000, with the possibility ofa lowingcommittees: Conservation,FieldTrips,Legislative,
special issuecomprisinganarticleentitled“SoilLichensandtheir Affairs,andPublicity. ContactJill Handwerkat(970)491
Application as Ecological Indicators in Sagebrush Habitats” con- 5857 ofAlice Guthrie at (303) 651-3127 for information
tributed by RogerRosentreter. and to volunteer.
New Business: The Rare Hant Book Fund was renamed the Bookreview:AquilegiaissolicitingareviewofTheLandscaping
MyrnaP. SteinkampMemorialFundandwill be used similarly Revolution: Garden withMotherNature, NotAgainstHer by
totheMarrFund.Interestgeneratedfromtheprinciplewill beded- Andyand SallyWasowski. Receiveafreecopyinreturnfora
icatedtotheprotectionofColoradorareplants,as well asresearch 500-1000wordreview.ContactLeoP. Bruederleat(303)861-
^'y^dvancingourknowledgeoftheseplants.TheBoardvotedtosplit [email protected].
Communications into twocommittees, one overseeingAquilegia Informationneeded:Wewouldliketocompilealistoffairs
publication and a second overseeing the CoNPS website, feo and shows atwhichtoeducatethe publicaboutCoNPS. If
Bruederle will serve as Chairofthe NewsletterCommittee, while you know of local or county shows that would be appro-
BobClarke will chairtheWeb Site Committee primarily oversee- priate, please call Alice Guthrie (303-651-3127) with the
ingContent. SaraHill, whow^ commendedforherworkas web- name,contactinformation,and general datesofthevenuel
master, will remain in that position. Jill Handwerk proposed a
cooperative effort with Colorado State University to bring Dr. AQUILEGIA Deadline Approaches
Gary Nabhan(Director,TheCenterforSustainableEnvironments, Submit on or before December 2pO0
1,
University ofNorthernArizona) toFortCollins inordertodeliver
a CSU Provost Lecture on EnvironmentalAchievement. The Please submit contributions forVol. 25 No. 1 ofAquilegia on or
Board approved allocating $5(X) toward an honorarium for Dr. beforeDecember 1,2(KX). Includeauthor’snameandaddress,and
Nabhan, who would also conduct a Society workshop, date and follow theformatfrom previous issues ofAquilegia as closely as
timeto beannounced. Inadiscussionofthedefinition oftheterm possible. Previously published articles submitted for reprinting
tnhaetifvoel,laoswiinngnaftriovmetphleanOtx,ftohredBoDiacitdioangarreye.d“tAoptpelmipeodratroilaysapcecceipets rseuqbumiirtetepdervmiiassei-omnaiflroormothnedeidsiktsor—ofMqrAigCinaplrepfuebrlaibclayt,ioonr.AIrBtiMcl—es
thatoccurs naturally in an areaad therefore one that has not been are much appreciated. Please submit the contribution as a MS
introduced by humans either accidentally or intentionally. Of WordorRTF(rich textformat) document.
plants found in a particular place, the term is applied to those
species that occur naturally in the region and at the site.” Gay The Importance of Native and Invasive
Austin will revise this definition based on the ensuing discussion. Species in Land Management
PeggyLyonpresentedFundingGuidelinestOTAdHocRequests, Society of Range ManagementAnnual Meeting
which was approved. The Board subsequently approved a request
for$500fromCityofBoulderOpenSpaceforrevisingandreprint- November 30 - December 1, 2000
ing the Russian olive brochure, on which the Society will be The Society for Range Management, USDA Forest Service,
le'^knowledged. LisaTaskerannouncedtheEcologicalRestoration BLM,ChaffeeCounty,ColoradoSeedGrowersAssociation,and
>^^Conference being sponsored by the Central Rockies Chapter of PawneeButtesSeedareputtingonaworkshopinFortCollinson
the Society for Ecological Restoration, for which Lisa hopes the invasive speciesandthe use ofnativeplants infevegetation. For
Board will considerlendingsupport. more information, pleasecontactDon Hijarat (970) 356-7002.
Page 4 Aquilegia Vol. 24 No. 5
CeRSER
NEWS Ecological Restoration Conference
SOCIETY
April 26-27, 2001
The Central Rockies Chapter of the Society for Ecological
Request For Proposals Restoration announces a very exciting regional conference:
^‘Restoring the Rockies: Restoration and Conservation
The John W. Marr Fund StrategiesintheWest,”tobeheldonThursdayandFriday,April
OnMarch24, 1990,the BoardofDirectorsofthe Colorado Native 26th and 27th, 2001 in Keystone, Colorado. With multiple orga-
Plant Society established the John W. nizationsthroughouttheRockiesnowpracticingrestorationwith-
Marr Fundfor the award of small grants inadiversityofecosystems,ameetingtoexchangestrategiesand
in support of research in plant biology. bring together a wide variety of individuals, groups, and coali-
The Fund honors the late Dr. John Marr, tions shouldbeofser\dcetoall. Inadditiontoexchangeofideas,
Professor at the University of Colorado the hope is that the conference will also help organizations form
and the first President of the Colorado alliances so as to better ensure the long-term survival of our
Native Plant Society. The Fund was diversehistoricalecosystemswithintheRockyMountainRegion.
establishedtoencourage,throughthepro- Keystone Resortis only 75 miles from Denverandmoreconve-
vision ofsmall grants-in-aid, research on niently locatedthantheFrontRangeforfolkscomingfrom west
the biology and natural history of ofthecontinental divide. Theresortclaimsthelargest,most ver-
Colorado native plants. Both field and satile meeting facilities in Colorado to say nothing of the sur-
laboratorystudiesareeligibleforfunding. roundings and recreational activities. Rates should be very rea-
Awards of $500 or less are typical and sonable,aslateApril islowseasonforKeystone. Don’tmissthis
may represent the sole support of a pro- exciting opportunity to find out more about the practice ofeco-
ject, may be part ofjoint funding agreed logical restoration in yourlocal ecosystems andhow itrelatesto
upoiiwithothergrantingorganizations,or the biggergoal ofconserving ournative plantcommunities.
maybeusedtosupplementotherfunding Submitting a PaperorPoster.
obtmned bytheinvestigator(s). Acall for papers is in progress.Abstracts and posters pertaining
Aquilegiachrysantha toall areasofrestorationarecurrently beingaccepted. Emphasis
Artist:JanWingate will be placed on riparian and wetland areas, rangeland restora-
P. Steinkamp Memorial Fund tion, restoration on private lands, restoration education (eg.,
On September22,2000, theBoardofDirectorsrenamed the Rare schoolyard habitats), restoration in rights-of-ways, and urban
Plant Book Fund to honorDr. Myma P. Steinkamp. Myma was a restoration. Proposals are welcome from all disciplines, back-
founding member of the Society, and serv^ many years both as grounds, and experience levels. Presentations should be
Treasurer and Chair of the Membership Committee. She also designed to be 20 minutes long, with an additional five minutes
served on the Conservation Committee, helping to establish the for question and answers. Longer time slots (up to 30 minutes)
Society’s role in Colorado plant conservation and public lands may be available if space permits. Please make the necessary
planning. She participated actively in the preparation of both edi- arrangements if your presentation does not fit the allotted 20
tions of the Society’s publication Rare Plants of Colorado. minutes. Duetotheanticipated diversity oftheaudience, papers
Although this fund had been specifically designated for publica- shouldincludeabriefbackground. Forposterpresentations,pre-
tionofRarePlantsofColorado,thescopeofthefundhasnowhas senters must provide their own free-standing bulletin boards.
been broadened to include support for research on rare Colorado Abstracts and proposals should begin with the name and orga-
native plants, through the provision of small grants-in-aid. Both nizational affiliation ofeach author with the presenting author’s
field and laboratory studies are eligible for funding. Awards of name underlined. Also include the address, phone and e-mail (if
$500orlessare expectedand may representthe sole support ofa available)ofthecorrespondingauthor,followedbyadescriptive
project,maybepartofjointfundingagreeduponwithothergrant- title and a text summary notexceeding 300 words. Submission
ing organizations, or may be used to supplement other funding deadline is December 1, 2000, although proposals may be
obtained by the investigator(s) for research in Colorado. accepted beyond thatdate on aspace available basis. Electronic
Contributionsto the memorial fund may bemade to: submissions as MS Word documents or RichText Files (rtf) are
preferred and should be submitted to Jody Nelson at
MymaR SteinkampMemorial Fund
[email protected]. Ifelectronic submissionis notpossible,
c/o Colorado Native Plant Society
hard copyordisk should be mailed to:
P.O.Box200
CentralRockiesChapteroftheSocietyforEcologicalRestoration
FortCollins, CO80522 P.O. Box 17644
The Board of Directors is now soliciting proposals for both the Boulder, Colorado 803040644
Marr Fund and Steinkamp Memorial fund, for which the dead-
Forthe most up to date information, please send e-mail to lisa-
linesareJanuary 15,2001. Informationonguidelinesand require- [email protected]. Find alinktothe Central Rockies Chapter
ments for proposals may be obtmned fiy contacting Board mem- through the international SER website www.ser.org.
berEric Laneat(303) 239-4182.
Vol. 24 No. 5 Aquilegia Page 5
MARR
FUND: Reports from Recipients
holistic Survey of South Central Colorado Status of the Green Sedge in Colorado
Brian Elliott Shannon Kuchel
During 1998 and 1999, an intense floristic survey was performed Carex viridula Michx. (Cyperaceae), the
in south central Colorado. This study, supported in part by the green sedg^, is a short-lived, graminoid
JohnMarrFundoftheColoradoNative PlantSociety, isthefocus perennial with adensely caespitose habit.
ofmyMaster'sThesisattheUniversityofWyoming. Fieldworkis It is monoecious, with a single terminal
now complete and has resulted in over 20,000 numbered collec- staminate spike and several sessile pistil-
A
tions, with detailed locationandecological dataforeach. setof late spikes. It typically occupies moist,
specimens will be retained at the Rocky Mountain Herbarium, early successional sites characterized by
with duplicates goingtootherpublic herbaria. fluctuatingandunpredictablewaterlevels.
The study area includes the Wet Mountains, Sangre de Cristo These sites vaiy from calcareous, acidic,
Range, Culebra Range, Spanish Peaks, and surrounding areas. sandy, or organic shorelines; runnels in
Ranging from the edge of Colorado's eastern plains up to an limestone barriers; wet meadows; marsh-
extensive alpine zone, the regionencompasses over5,000 square es; on borders of streams, ponds, and
miles, with a rich diversity of vegetation types. The area has lakes;tofens. Carexviridulaisdistributed
received relativelylittleattentionfrombotanists previouslyand is throughout northern and central North
hardly mentioned in Weber and Wittman's Colorado Flora: America. It also extends to the southern
EasternSlope. Rocky Mountain region in several dis-
junct sites, including alpine wetlands in
Thesurveydocumentedapproximately 1150taxa.Thisincludes44 Colorado, where it is rare. Although C.
taxafrom the ColoradoNatural Heritage Program’s listofspecies viridula has been assigned a global rank-
of spiecial concern. Notable collections include dwarf milkweed
ing ofG5, indicating that itis demonstra-
{Asclepias uncialis), Colorado larkspur {Delphinium alpestre), bly secure globally, it has received astate Artist;JanWingate
Gray's Peak whitiowgrass {Draha grayana). Smith whitlowgrass
rankingof SI, indicating that itis critically imperiled inColorado.
.smithii),show}'prairieGentian {Eustomagrandiflorum),spiny The purpose of this research was to describe genetic diversity in
^ster {Herrickia horrida)., 12 populations of rock-loving
populations of C. viridula from Colorado, and determine if these
Neopanya {Neoparrya lithophila),alpine poppy {Papaverradica - populations exhibited reduced genetic diversity when compared to
turn ssp. kluanensis), pictureieaf wintergreen (Pyrola picta), and otherpopulations ofthis speciesfrom NorthAmericaandEurope.
Altai stitchwort {Stellaria
irrigua). Also collected were In ordertoquantify geneticdiversity, small samples ofleaftissue
Cymopterus constancei, were collected from 350 individuals representing seven extant
newly described by Dr. Ron populations ofC. viridula in Coloradoand 179individuals repre-
Hartman inBrittonia., andthe senting eight populationsfrom elsewhere throughout the rangeof
soon to be published this species in NorthAmerica. Starch gel electrophoresis ofsolu-
Oonopsispuebloensis. ble enzymatic proteins coupled with substrate specific staining
wasusedtodeterminetheamountanddistributionofgeneticvari-
This study is only part of a ation in these populations. These data were compared with data
larger Flora of the Rocky
previously reportedforEuropean populations ofthis species.
Mountains Project being
conducted by Dr. Hartman, Usingthesetechniques,nogeneticvariation,eitherwithinoramong
Curator of the Rocky populations from Colorado or elsewhere in NorthAmerican, was
Mountain Herbarium at the detected. However, NorthAmerican populations were genetically
University of Wyoming differentiated from European populations, with significantly more
(http://www.rmh.uwyo.edu). diversity maintained by the latter. The surprising lack of genetic
Many thanks are due to the diversity in NorthAmerican populations is probably thecombined
Colorado Native Plant resultofhigh levelsofselling and inbreeding, restrictedecological
Society (andespecially tothe amplitude, and genetic drift. Genetic bottlenecks are presumed to
supporters of the John Marr have occurred as a result of climate changes associated with
Fund), as well as to Nancy Pleistocene glaciationorfoundingevents associated with coloniza-
Ryke of the San Isabel tionofNorthAmericabyproposedancestral Europeanpopulations.
tionalForestforgenerous- I would like to thank Dr. Leo R Bruederle for his guidance
^Vsupportingthis research. throughout this project. I am also grateful for support from the
Eustomagrandiflorum Colorado Native Plant Society, Sigma Xi, and the Department of
Artist:JanA^ngate Biology atthe University ofColoradoatDenver.
Page 6 Aquilegia Vol. 24 No. 5
CHAPTER NEWS
Boulder Chapter Storage Laboratory (NSSL) onthecampus is primarily montane, with both lowerand
ofCStJ. The NSSL is locatedjust west of upper montane forest well-represented.
Monthly meetings are held October the railroad tracks, about one block north Leo P. Brnederle (Associate Professor,
throughAprilattheFoothillsNatureCenter, of Pitkin Street (enter on the west side of CU-Denver) will discuss ongoingresearch
locatedinBoulderat4201 NorthBroadway, the building). For more information, con- conducted along with CoNPS members
opposite its intersection with Sumac Ave. tactChapterPresidentDonHazleltat(970) SaraE. HillandKevinFehlberg,theobjec-
The Chapter now meets on the second 834-1493. tiveofwhichwastocircumscribetheflora
Wednesday of each month, at 7:00 pm for ofthe RampartRange.
business,aspeaker,booksales,andsocializ- November 9, 7:00pm
ingWithrefreshments. Pleasejoinusat5:00 Weed Warrioring and other Botanical January 30, 7:30PM
PM for supper with our speaker at Efrain’s ExploitsatDinosaurNationalMonument New Species in the Myrtle Family from
Mexican Restaurant, 1630 N. 63rd St, just Tamara Naumann (Botanist, Dinosaur the Rich Flora ofQueensland,Australia
north of Arapahoe in Boulder. For more National Monument) will repeat her Queensland is mega-rich in its biodiversi-
information, contact Chapter President November8 BoulderChapterProgram. ty,asevidencedbyits9,000speciesofvas-
Caiyl Shieldsat(303)665-6312orProgram cular plants. Many new species are
ChairCarolynCrawford(303)665-6903. December5, 7:00pm described annually, particularly from the
Ve^tableMotor-oilforColoradoVehicles
more tropical areas of the state. Recent
November8j7:00PM Duane Johnson (Professor ofAgriculture, research has focused on describing new
Weed Warrioring and other Botanical CSU) will discuss his extensive work with
species and taxonomicallyrevising several
ExploitsatDinosaurNationalMonument leSserknownColoradocropsandplantprod- genera of fleshy-fruited members of the
Tamara Naumann (Botanist, Dinosaur ucts. He will also review informationon the Myrtlefamily(Myrtaceae). Dr, NeilSnow
National Monument) will present some of toxicityofBTcompollentonativeinsects. (Assistant Professor, UNC) will provide a
htheerMexopneurmieenncte,sionvcelrudtihneglatshtosseevdeeraalliynegarwsitaht FTeobbreuaarnyno6u,n7c:e00dpm bwritohada poavretrivcuileawrfoofcutshoenAQuuseteranlsilaannd.flora,
weeds,rareplants, tourists,and ranchers.
December 13,7:00pm Metro-Denver Chapter Plateau Chapter
The World of Wildland Fires, Indoor monthly meetings are held at7:30 Workshops, field trips, and other activities
PrescribedBorns,andFireEcology PM from September through April in the are scheduled throughout the year. For
Coloradojustexperienced one 6fits worst Morrison Center at the Denver Botanic more information, please contact Chapter
years for wildland (forest, brush, and Garden. For more information, contact PresidentJeanneWengerat(970)256-9227
range) fires. Justin Dombrowski (Fire PresidentDeniseLarsonat(303)733-4338 or by e-mail at [email protected] or
Management Officer, Boulder Fire or Leo P. Bruederle at (303) 861-4119 or ProgramChairLori Brummerat(970)641-
Department) is involved in many facets of bye-mail at [email protected]. 3561 [email protected],
wildland fire management throughout the
fireseasonandacrosstheState. Hewilldis- October31, 7:30 pm November 18, 10:30am
cuss his experiencesduringthe pastyear. StatusoftheGreenSedgein Colorado Plateau Chapter Annual Planning
The green sedge, Carex viridula Meeting and Potluck
January 10,7:00pm (Cyperaceae), occurs in wetland habitats Please note this change in date!
TheRocky MountainArsenal scattered throughout northern and central Following our organizational meeting
The RockyMountainArsenal wasformer- NorthAmericaandEurasia. Itsdistribution (workshops, field trips, and elections) and
ly involved in weapons work during the alsoextends to several disjunct sites in the lunch, we’ll have a seed and catalog
Cold War. Since being decommissioned, alpine wetlands of Colorado, where it is exchange. Members will then have an
much of the area has been restored as a critically imperiled. Shannon Kuchel, opportunity to show off some ofthe great
wildlife refuge. Suzanne O’Neill 1999 recipient of a John W. Maix Fund slides they took over the year. Dust your
(President and Director, Rocky Mountain research grant, will discuss her research cameras offand buy some slide film! This
Arsenal Wildlife Society) will discuss the addressing genetic diversity maintainedby should be fun! Meet at the Forest Service
changes that have occurred over the years Colorado populations ofthe green sedge. building in Delta.
andthefutureofthe site.
December5,7:30 pm Southwest Chapter
Fort Collins Chapter Floraofthe RampartRange.
The Rampart Range is one ofa numberof For Chapter news and activities, pleasf
Indoor monthly meetings are scheduled distinct geologic features comprising the contact Chapter President Sandy Friedle^L:::;?^
through April at 7:00 PM in the Main Front Range of the southern Rockies in at (970) 884-9245 or by e-mail at fried-
Conference Room of the National Seed Colorado. The vegetation ofthe Ramparts [email protected].
.
Vol. 24 No. 5 Aquilegia Page 7
Colorado Native BoardofDirectors
Plant Society Jill Handwerk(01) . . FortCollins. . 970-491-5857
BillJennings(01). . .Louisville . ..303-666-8348
AndyKratz(01)... .Lakewood. . .303-914-8291
PatPloegsma(01). . . Strasburg. . . . 303-622-9439
Aquilegia is published four to six times per Ericyne(01) Lakewood. . . 303-239-4182
year by the Colorado Native Plant Society. DavidAnderson(02) FortCollins. . 970-484-0774
This newsletteris available to members ofthe GayAustin(02) .... Gunnison. . . . 970-641-6264
Society and toothers with aninterestin native KathyDarrpw(02). . CrestedButte. 970-349-0743
The Colorado Native Plant Society is a non- plants. Contact the Society for subscription GwenKittel(02).. . .Boulder 303-258-0908
profit organization dedicated to the apprecia- information. Articles for Aquilegia may be Laurel Potts(02). ...FortCollins. . 970-484-4076
tion and conservation of the Colorado native used by other native plant societies or non- Neil Snow(02) Greeley 970-330-4823
flora. Membershipis open toall with an inter- profitgroups iffullycitedtoauthorandattrib- ChapterPresidents
est in our native plants, and is composed of utedtoAquilegia.
Boulder.......Caryl Shields ....303-665-6312
plant enthusiasts both professional and non-
professional. Pleasedirectallcontributionstothenewsletterto; FortCollins....DonHazlett ..... 970-834-1493
LeoP.Bruederie Metro-Denver . . DeniseLarson . . .303-733-4338
Pleasejoinus in helping to encourage interest Biology,Campus Box 171 Plateau JeanneWenger. . .970-256-9227
in enjoying and protecting Colorado’s native University ofColoradoatDenver Southwest SandyFriedley. .1970-884-9245
plants. The Society sponsors field trips, work- P.O.Box 173364
shops, ahd otheractivities throughlocal chap- Denver,CO 80217-3364 Committees
Conservation...VACANT
tersandstatewide. ContacttheSociety,achap- E-Mail: [email protected]
terrepresentative,orcornmitteechairformore Education .....TinaJones 303-794-2647
information. Shortitems suchas unusual informationabout andCo-chairVACANT
aplant,alittle known botanical term,etc.,are FieldStudies .. . LoraineYeatts . . .303-279-3427
ScheduleofMembWshipFees especially welcome. Previouslypublishedarti- FieldTrips.....VACANT
ySufpeporting : . . . . ,. . . .. ..$.2$5500 cslioens.sCuabmmeitrtae-drefaodryrelpirneintaritngorreoqtuhierer iplelrumstirsa-- HoRretsitcourlattuiroenan.d...KMadthLyisDaaTmaasske.r.....390730--554443--31643932
Organization . . . . .$30 tions are also solicited. Pleaseinclude author's Legislative
FamilyorDual .$15 nameandaddress,althoughanonymitymaybe Affairs ..VACANT
. .
ndividual .$12 requested. Articles submitted via e-mail or on Membership. . . .EricLane..... . 303-239-4182
IStudentorSenior .$8 disks(MACpreferably,orIBM)areappreciat- andSueMartin. . . 970-226-3371
ed. Please indicate word processing software Newsletter LeoP.Bruederie. .303-556-3419
MembershipRene^al/Information andversion;ifpossible,submitasanRTF(rich Publications....JeffDawson 303-722-6758
Please direct all membership applications, textformat) file. andVelmaRichards303-794-5432
renewals, and address changes to the Publicity VACANT
Membership Chairperson, Colorado Native Officers RarePlant ... Eleanor
Plant Society, P.O. Box 200, Fort Collins, CO President JillHandwerk . .970-491-5857 Monograp.h. . . . VonBargen ....303-756-1400
80522.Heasedirectallotherinquiriesreg^ding Vice-President. . .DavidAnderson. 970-484-0774 WebSite BobQarke 970-242-6067
theSocietytotheSecretaryatthesameaddress. Secretary AliceGuthrie. . .303-651-3127 Workshops BillJennings ....303-666-8348
Treasurer DeniseCulver . .970-225-1930
MEMBERSHIP APPLICATION FORM
Name(s) MEMBERSHIPCLASS
(Selectone)
Address ' Individual, $12.00
^
Family/dual, $15.00
(Address) Senior/retired, $8.00
Student, $8.00
City - State Zip Corporate, $30.00
Supporting, $50.00
Phone Li 1 E-mail .
Chapter; Boulder ____ Fort Collins Metro Denver Plateau Southwest
Inadditionto rny membership, I have included $ as a contribution to theJohn Man-
Fund (endowment in supportofsmall grants-in-aid ofresearch), $ as acontribution to
^.le MymaP. Steinkamp Memprial Fund (endowmentin support ofsmaH grants-in-aid of
research), or $ as a general contributionto the Society.
CONPSISA NON-PROFITORGANIZATION -DUESAND CONTRIBUTIONSARE TAX-DEDUCTIBLE
CHAPTER MEETINGS AND EVENTS Plateau Chapter (continued)
Boulder Chapter January 27 Worikshop: How to know the Families
ofthe Western Slope
November8 WeedWarrioringandotherBotanical
ExploitsatDinosaurNat’l. Monument March 17 Workshop: Grasses ofthe Western
Slope
December 13 TheWorld ofWildland Fires,
Prescribed Bums, and Fire Ecology SOCIETY EVENTS
January 10 The Rocky MountainArsenal Board Meetings
Fort Collins Chapter November4 Longmont, Colorado
November9 WeedWarrioringandotherBotanical Workshops
ExploitsatDinosaurNat’l. Monument November4, 5 How to Know the Families
December5 VegetableMotor-oilforColoradoVehicles First session: Saturday
Second session: Sunday
Metro-Denver Chapter
December9, 10 Small Plant Families
October31 Statusofthe Green Sedge in Colorado
First session: Saturday
December5 FloristicAnalysisofthe Rampart Range Second session: Sunday
January 30 New Species in the MyrtleFamily from January 13, 14 Grasses: TheTribe Hordeae
the Rich FloraofQueensland,Australia First session: Saturday
Second session: Sunday
Plateau Chapter
November 18 Plateau ChapterAnnual Planning Febraary 10, 11 Cirsium: The Native Thistles
Firstsession: Saturday
Meeting and Potluck
Second session: Sunday
ColoradoNativePlantSociety
P.O. Box 200
FortCollins,Colorado80522
http://www.conps.org
TIME SENSITIVE MATERIAL