Elytra

2006·168.1 MB·

CyCh「iniandCarabinifromcentralchina
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242 Ytiki IMURA
2-2. LeptinocarabuszhubaJ'ieshunwalrtgping IMURA,subsp
nov.
(Figs3,11)
Description. Length (includingmandibles): 18.6-20.4mm,早, 20.0-22.4mm.
,
Externalappearancealmostasinsubsp.laoJunensis,butapicallobeofaedeagusrobuster
andmorestronglybent right lateradinbothdorsal andventral views.
Typeseries. Holotype: ?,SEslopeofShunwangping [舜王坪],2,040-2,200min
alt.,35°24'N/111°58'E,onMt.LiShan [1?1;山],onthebordersofYichengXian [翼城
], Qinshui Xian [沁水 ] and Yuanqu Xian [垣曲一 ] ofsouthernShanxi, Central
China, l3-V~4-VII-2006,preservedintheDepartmentofZoology,NationalScience
Museum (Nat. Hist), Tokyo(=NSMT). Paratypes:15 33早早,samedataasfor
,
theholotype,preserved in thecollectionsof Y.IMURA& J. TURNA.
3 TitanocarabussuineixiangensisIMURA,subsp
nov
(Figs 4,12)
Descrpptt1lon. Length(includingmandibles):d'',32.6-35.8mm,早,36.1mm. Simi-
lartothenominotypica1subspecies,butdistinguishablefromitbyhavingmorestrongly
protruded pronota1 hind angles, less remarkably meandered striae between elytra1
intervalsandmorestronglyconvexright lateralmarginofaedeagusbeforeapex.
Typeseries. Holotype: Baotianman Nature Reserve, l,500-1,750m in alt.,
,
3°331'N/111°56'E,ontheFu'niuShanMountains,inNeixiangXian,ofwesternHenan,
CentralChina,10-V~6-VII-2006,preservedinNSMT. Paratypes: l ,1早,samedata
asfor theholotype,preservedinthecollectionsofY. IMURA(早) &J.TURNA( ).
4 Piocarabus(Qinlingocarabus) kitawakianusshirenshanicusIMuRA,subsp
n ov
(Figs 5,14)
.I)escription. Length (includingmandibles): ,22.3-25.4mm,早,22.7-26.5mm.
Distinguishable from thenominotypica1subspecies (IMuRA, 1993) in the following
respects: l) sizea littlesmaller; 2) lateralsidesof pronotum less remarkablysinuate
Figs9-18. MalegenitalorganofnewlydescribedsubspeciesandtheiralliedtaxafromChina. -
s 9,Cychrusblsp加osus1/ena川cus(a,rightlateralview;b,dorsalview);10, ept加ocarab1e 加z baJl
1aoJunetlsis(a,rightlateralview;b,rightsubventralview);11.L.z.shtm、,angping(a,rightlateral
view;b,rightsubventralview);12,Tita1locarabussl1l,letxlangensls(a,right lateralview,b,apical
part in ventral view); 13, T. s.stii from Beijing(a, rightlateral view, b apical partinventral
view); l4, Piocarab1ls (gl'll加gocara加s) 1lawak1a川Is s1/1rens11a川ells (a, right lateral view; b.
apical part in rightsubdorsal view;c,dittoindorsal view;d,apicalpartofendopha11us);15,
Eucarabus (Parhomoptertls)manifest11s lis11atlensls(a, rightsubdorsal view;b, right lateralview;
c, dorsal view: d, endophallus viewed from inflexed side; e, ditto in right lateral view); 16,
Pagocarabuscrassesculptuscrassesc111ptusfromBeijing(apicalpartofaedeagusinrightsubdorsal
view);17,P.c.wan)cianshanus(a,rightlateralview;b,apicalpartofaedeagusinrightsubdorsal
view;c,dorsalview);18,P.c. ll,nontaluls(apicalpartofaedeagusinrightsubdorsalview).
CychriniandCarabini fromCentralChina 243
1
, 10-a
j
l4●-
14-b
a
15-e
,
. 17-c
l5-c
l
244 Yuki IMURA
beforehindangles;3)hindanglesofpronotumshorterandalittlelesssharplypointed
at tips;4) aedeagusmuchslenderer,withthemedianportionnarrower, right lateral
marginlessstronglyconvexlateradatmiddleandmoreweaklyconcavetherefromtothe
apex, apical lobe longer andslenderer; 5) alargeU-shapedpigmentedplateon the
ventralwallofendopha11usbeforelacinianarrowerandsmaller. Fromsubsp huodi-
tangicus(CAvAzzUTI, 1999), thenewsubspecies isdistinguishedbywider pronotum
with robuster hindangles,more remarkably unevenpronota1disc, longer and less
stronglybent aedeagalapex, etc.
, Mt.ShirenShan, 2,000minaltitude, 3°342'N/11°2
Typeseries. Holotype:
15'E,ontheFu'niuShanMountains,onthebordersofSongXian, LushanXianand
Nanzhao Xian of western Henan, Central China, 9-V~9-VII-2006, preserved in
NSMT. Paratypes:6 8早早,samedataasfor theholotype,separatelypreservedin
,
the collections of Y. IMURA & J. TURNA.
5. SmoleptocarabusmarcithaciromanellusIMURA, 1999
Carabus (」Leptocarabus)ma・tci111acl・,omaniIMURA, l998,Jpn.J.syst.Ent.,Matsuyama,4,p 44,figs 6,17;
typelocality:nearthebordersofShaanxiandHenan(33°48'-53'N/110°40'-46'E),China. NeeCarabus
(Chrysocarabus)plmctatoatl,'attls''omantRAYNAUD,1968,Bull.Soc.finn.Lyon,37,p 303.
Carabus(Leptocarabus)marcilhac1romat1e11usIMURA,1999,Jpn.J.syst.Ent.,Matsuyama,5,p.143.
Specimensexamined. 10早早, Mt. ShirenShan, 2,000m in altitude, 33°42'N/
112°15'E,ontheFu'niuShanMountains,onthebordersofSongXian, LushanXian
and NanzhaoXianof westernHenan, CentralChina, 9-V~9-VII-2006, preserved in
thecollections of Y. IMURA & J. TURNA.
Notes. SubspecificidentificationoftheShirenShanspecimensisuncertain,sinceI
wasunabletoexaminethemaleanditsgenitalorgan. SofarasIhaveexaminedthe
femalespecimens,however, it isalmostidenticalwiththosefromBaotianman,andcould
be identified as subsp romane11us described from near the borders of Shaanxi and
Henan.
6. Scambocarabusshaan;c'iensisDEuvE,1991
Carabus(Tomocarabus)s11aanx::1enslsDEUvE,1991,Nouv.RevueEnt.,Paris,(N.S),8,p ie5:typelocality:
Chine,Shaanxi,AnkangXian.
Specimensexammed. 20 ,25早早,Mt.ShirenShan,2,000minaltitude,33°42'
N/112°15'E,ontheFu'niuShanMountains,onthebordersofSongXian,LushanXian
andNanzhaoXianofwestern Henan,CentralChina,9-V~9-VII-2006,preservedin
the collections of Y. IMURA & J. TURNA.
7.ScambocarabussculptipennisCHAUDoIR,1877
CarabussctltptppennisCHAUDolR,1877,Dt ent.Z.,21,p 75;typearea:“IenerddelaChine'
CychriniandCarabini fromCentralChina 245
Specimensexamined. 2早早,SE slopeofShunwangping, 2,040-2,200m inalt.,
35°24'N/111°58'E,onMt.LiShan,onthebordersofYichengXian,QinshuiXianand
YuanquXianofsouthernShanxi,CentralChina,13-V~4-VII-2006,preservedinthe
collection of J. TURNA.
8. CarabusgranulatustellurisBATES,1883
Carabusgran1llatusvar.te111lrtsBATES,1883,Trans ent.Soc. Lend.、l883.p 223;typelocality:Tonosawa,
central Honshu.
Specimensexamined. 15 ,9早早, Mt. WanxianShan [万仙山], 840m in alt.,
35°42'N/113°36'E,inXinxiangShi [新 市] ofnorthernHenan,CentralChina,15-V~
1-VII-2006,preservedin thecollectionsofY.IMURA&J.TURNA.
Notes. TheWanxianShanpopulation issomewhatcharacteristicinhavingdark
bluish coloration on the upper surfaceof thebody. Otherwise, however, it is not
remarkablydifferent fromthosedistributedinnortheasternChina,northernKoreaand
Honshu,Japan.
9-1. Eucarablis(Parhomopterus) manifeistusmanifestus KRAATz,1881
Carabustnanifest1lsKRAATz,1881,Dt ent.Z..25,p 268: typelocality:GebirgenwestlichyonPeking.
Specimene;x;amined. 1早,Mt.WanxianShan,840minalt.,35°42'N/113°36'E,in
XinxiangShiof northernHenan, CentralChina,15-V~1-VII-2006, preservedin the
collectionof J. TURNA.
9-2. Eucarabus (Parhomopterus)manifeistuslishanensisIMuRA,subsp nov.
(Figs 6.15)
Description. Length (includingmandibles): ?,22.3-22.4mm,早,22.0-23.5mm.
Bodyabovedark coppery brown. Differs from thenominotypica1subspecies in the
following points: 1) pronotumwith the front anglesa little less roundly protruded
anteriad and the hind angles shorter; 2) elytraa little robuster; 3) aedeagus more
remarkably inflatedventradat apical third, with theapical lobewiderand longer in
lateral view, anda little lessstronglybent right laterad indorsal view. Fromsubsp
guanmenshanus (IMuRA, 1991,p 274),thenewsubspeciesisreadilydiscriminatedby
differently shaped pronotum and aedeagus, as well as differently sculptured elytra1
surface.
Typeseries. Holotype: ,SEslopeofShunwangping,2,040-2,200minalt.,35°
24'N/111°58'E,onMt. LiShan, onthebordersofYichengXian,Qinshui Xianand
YuanquXianofsouthernShanxi, Central China, 13-V~4-VII-2006,preservedin the
collectionoftheDepartmentofZoology,NationalScienceMuseum(Nat.Hist),Tokyo
(=NSMT). Paratypes: 3 5早早,samedataas for theholotype, preserved in the
,
246 Yuki IMllRA
collectionsofY.IMURA(早)&J.TURNA( )
10-1. Pagocarabuscrassesculptuscrassethoracis KLEINFELD,1999
(Fig.16)
Carabus(Pagocarabus)classethoraclsKLEINFELD.1999,Ent.Z.,Essen,109,p. l4,Abb.13,16;typelocality:
China,Henan,LaojunMt. (33.43/111.37),Luangchuan [sic] (misspellingofLuanchuan).
Carabus(Pagocarabus)crassesculptuscrassethoracls:Deuve,2004,111.Cat.Ca・labusWorld,Pensoft,Sofia/
Moscow,p 279.
Pagocarabuscrassesculptuscrassethoraels: IMURA,2005,Elytra,Tokyo,33,p 669.
Specimensexamined. 13 , 12早早, Mt.Shiren Shan, 2,000m inaltitude, 33°
42'N/11°215'E,ontheFu'niuShanMountains,onthebordersofSongXian,Lushan
Xian and Nanzhao Xian of western Henan, Central China, 9-V~9-VII-2006, pre-
served in the collections of Y. IMURA & J. TURNA.
l0-2. Pagocarabuscrassesculptuswan;x:1anshanusIMURA,subsp
n ov.
(Figs7,17)
Description. Length (includingmandibles): , 24.5-27.3mm,早, 27.4-29.0mm.
Alliedtothenominotypica1subspecies,but differs fromit in the followingrespects:1)
sizeapparently larger onanaverage; 2) pronotummuch wider, with thehindangles
hardlyprotrudedposteriad;3)elytra1costaelessfrequentlyconfluent withoneanother;
4) apical lobeof aedeagus1ongerand lessacutelynarrowed towardapex insubdorsal
view.
Typeseries. Holotype:?,Mt.WanxianShan,840minalt.,35°42'N/113°36'E,in
XinxiangShinorthernHenan,CentralChina,15-V~1-VII-2006,preservedinNSMT.
Paratypes:13 18早早,samedataasfor theholotype,preservedinthecollectionsof
,
Y. IMURA & J. TURNA.
10-3. Pagocarab1lscrassesculptuslimontanusIMURA,subsp nov.
(Figs 8,l8)
Description. Length (includingmandibles): ,22.1-24.9mm,早,24.4-25.5mm.
Most closelyallied to thenominotypica1subspecies, but distinguishable fromit in the
followingpoints: 1) colorationof dorsalsurfacemorebrightlygreenish blue inmost
specimens;2) pronotumalittlewider,withthelateralsidesalittlemoreremarkably
sinuate,andhindanglesmoresharplypointedattips;3)elytraalittlerobuster,withthe
striaebetween intervalsdeepersothat thecostaeof intervalsless frequentlyconfluent
with oneanother; 4) apical lobeof aedeagus alittle robuster and less remarkably
narrowed towardsapex insubdorsalview.
Type series. Holotype: (:i7', SE slope of Shunwangping, 2,040-2,200m in alt.,
35°24'N/111°58'E,onMt. LiShan,onthebordersofYichengXian,QinshuiXianand
CychriniandCarabini fromCentralChina 247
Yuanqu Xian of southern Shanxi, Central China, l3-V~4-VII-2006, preserved in
NSMT. Paratypes: 32 29早早,samedataas for theholotype, preserved in the
,
collection of Y. IMURA & J. TURNA.
ll. Cathaicusbrandti FALDERMANN,1835
CarabusbrandtiFALDERMANN,1835,Mem.Acad.Sci.St.Petersb.,2,p 338;typearea:“NChina”
Specimensexamined. 2 ,Mt.WanxianShan,840minalt.,35°42'N/113°36'E,
inXinxiangShiofnorthernHenan,CentralChina,15-V~1-VII-2006,preservedinthe
collection of J. TURNA.
12. CoptolabrussmaragdinuszhongtiaoshanusIMURAetYAMAYA,1994
Carabus (Coptolabrus)smaragdinuszhongtlaoshantls IMURA et YAMAYA, 1994, Gekkan-Mushi, Tokyo,
(275),p.14; typelocality:Mt.ZhongtiaoShan,ShanxiProvince,CentralChina.
Specimensexamined. 70 ,39早早,Mt.WanxianShan,840minalt.,3°542'N/
113°36'E, in Xinxiang Shi of northern Henan, Central China, 15-V~1-VII-2006,
preservedinthecollectionsofY. IMURA&J.TURNA.
要 約
ｾ 井付有希: 中国河南省と山西省におけるセダカオサムシ方と ォサ €シ族の記録 . - 中国
南省西部伏牛山塊東部の石人山, 同省;lヒ部の万仙山, および山西省南部のノl j山からセダカオサム
シ族1種とォサムシ族11種を記録し, これらのうちななつを新亜種として記載した. また, 伏牛
山塊の主峰, 老君山からlc,?'のみの標本に基づいて記載された“Carabus(Rliigocarabus)”1aoJunen-
sisを, LeptinocarabuszhubaJieの1亜?重へと降格した. 本種は当初, ドウガネオサムシ類に属する
独立種として記載されたが, 今回, 同山i?東部の石人山からあらたに得られた複数の此能雄を, (f
交尾器内装の形態まで含めて詳細に検討したところ, 力ザリオサムシ類のなかのウェヌストゥス
カザリオサムシ属Leptinocarabusに属するL.zhubaJieの1亜種とみなすべきであることがあきら
かになった.
References
BATES,H.W.,1883.SupplementtothegeodephagousColeopteraofJapan,chieflyfromthecollectionofMr.
GeorgeLEWIS,madeduringhissecondvisit,fromFebruary,1880,toSeptember,1881. 「ransent.Soc.
Lend.,1883:205-290,1pl.
CAvAzzUTl, P., 1999. SestocontributealiaconoscenzadelCarabus L delta Cina. Descrizione di nuove
specieesottospeciesdelleShaanxi,SichuaneYunnan(Coleoptera,Carabidae). Coleopteres,Guyancourt,
5: 115-137.
CHAUDolR, ru le Baronde,1877. NotesurquelquesespecesdoCarabesplatsduCaucase. Dt ent. Z, 21:
69-95.
DEUvE, Th., 1989. Carabidae et Trechidae nouveaux dos collectionsentomologiquesde la North-West
248 Yuki IMURA
AgriculturalUniversitydoYangling,Shaanxi (Coleoptera). Ento,notaxonolnIa,11:227-235.
DEUVE, Th., 1991. NouveauxCarabusdescollectionsdel'InstituteZoo1ogiquedePekin (Coleoptera,
Carabidae). Nottv. RevtleElit.,Pa・,ts, (N.S),8:101-108.
- 2004. IllustratedCatalogueoftheGenusCarabusoftheWorld(Coleoptera:Carabidae). 461pp.
Pensoft,Sofia/Moscow.
FALDERMA,、'N. F., 1835. Coleopterorumab111.BungleinChinaboreale,MongoliaetmontibusAltaicus
collectorum nee non ab111. TurczaninoffoetStschukinoeprovincial Irkutskmissorum illustrations.
Mom.Acad.Set.St.Petersb.,2:335-464.
IMuRA, Y.,1991. Notesoncarabidbeetles(Coleoptera,Carabidae)fromEastLiaoning,NortheastChina.
Elytra,Tokyo,19:273-283.
- 1993. New or littleknown CarabusandCyci1,-us (Coleoptera, Carabidae) from theQinling
MountainsinShaanxiProvince,CentralChina. Ibid.,21:363-377.
- 1993. A new Oreocarabus (Coleoptera, Carabidae) from theQinlingMountainsinShaanxi
Province,CentralChina. Ibid., 21:379-382.
- 1995. Neworleastknowncarabidbeetles(Coleoptera,Carabidae)fromtheDabashanMountains
atthenortheasternendofSichuanProvince,CentralChina. Ibid.,23:119-128.
- 1998. DescriptionsoftwelvenewCarabltsandCycllr11s(Coleoptera,Carabidae)fromChina.Jpn.
J.syst. Ent., Matsuyama, 4:39-49.
- 1999.TennewtaxaofthegeneraCarabusandCychrus(Coleoptera,Carabidae)fromChina. Ibid.,
5:135-144.
- 2002. Classification o「 the subtribe Carabina (Coleoptera, Carabidae) based on molecular
phylogeny. E:1ytra,Tokyo,30:1-28.
- 2005. CalosominaandCarabina(Coleoptera,Carabidae)fromtheBaotianmanNatureReservein
western Henan,Central China. Ibid.,33:665-670.
- &S.YAMAYA,1994. TwonewsubspeciesofCoptolabrtls(Coleoptera,Carabidae)fromShaanxi
andShanxi.China.Gekken-Mtishl,Tokyo,(275):12-24. (InJapanese,withEnglishtitle,descriptionand
summary.)
- & H.-Z.ZHOu, 1998. Ane、vTltanocarabus(Coleoptera,Carabidae) discoveredfromBeijing,
China. Elyt・,a, Tokyo,26:113-ll4.
& Z.-H.SU,1999. AnewspeciesofApotomopte''itsandanewsubspeciesofCyc11tus
(Coleoptera.Carabidae)fromChina. Ibtd.,27:651-654.
KLEINFELD,F.,1999. NeueTaxaderGattungCarabus(Coptolabrils,Apoto1r1opterus,Pagocarabus,Rhlgo_
carabus)ausdenchinesischenProvinzenHenan,HubeiundSichuan(Coleoptera:Carabidae:Carabini).
Ent. Z,Essen,109:1-48.
KRAATZ,G..1881. FiinfneuechinesischeCarabus. I)t elf. Z,25:265-269.
Elytra.T1okyo,34(2):249-256.November18.2006
Occurrenceof Yarnautidius (Coleoptera, Trechinae) in the
UpperHypogeanZoneofSouthwesternShikoku,
Southwest Japan
Shun-lobi UENo
DepartmentofZoology,NationalScienceMuseum(Nat.Hist)
3-23-l Hyakunin-cho,Shinjuku,Tokyo,169-0073Japan
and
Takao NAITo
3-4-13Ikeda,Neyagawa-shi,Osaka,572-0039Japan
Abstract Twonewspeciesoft1、etrechinegenusYa,nautldiusaredescribedfor
the first time from the upper hypogeanzoneofsouthwesternShikoku,Southwest
Japan. Bothseemtobelongtothe11ozlm1lcomplexofthegroupof「.fatipe11川s,and
arereadilydistinguishedfromY. tloz1l,nt1andfromeachotherbypeculiaritiesof the
aedeaga1apical lobe and of the copulatory sclerite. The new namesgiven are
Yamautidius (s.str)crosonand Y. .silo'no.
Twenty-four species of small delicate trechinebeetles belonging to thegenus
Yalmautidiushavebeen knownuptothepresentonly from limestonecavesandmine
aditsinthewesternpartof theIslandofShikoku,SouthwestJapan (cf.UtNo,1957,
1960, l978,1982,1990). Theyareclassifiedintotwosubgenera,Yamautidius(s.str)
andMiyamaidius(UtNo,1978),andthemembersoftheformerarearrangedintonine
complexesof threespecies-groups(cf.U直NO,1982,pp 59-61,1990). Occurrenceof
upperhypogeanpopulationshaslongbeenanticipated,butourexpectationhasnotbeen
fulfilleduntil thelastspring. Anotherexpectationunfulfilledisclarificationofsouth-
westward extension of the generic range; thepreviouslymarked southern limit of
distributionisArinoki-no-kanaudoMineinTowa-mura, locatedatthenorthernsideof
theuppermaincourseoftheShimanto-gawaRiver,whichisinhabitedbyY.uozumiiS.
UtNo(1982,pp.13,30,figs 24-26;cf.No8onfig55,inU直No,1982,p 63). Towards
thesouthwest of Towa-mura, thereisa fairlywideareadominatedby theUwaHills,
surroundedbythemiddlecourseoftheShimanto-gawaRiveranditsmaintributary,the
Yoshino-gawaontheeastandnortheast,bytheNakasujiDepressiononthesouth,and
bytheBungoChannelonthewest. Noanophthalmictrechinebeetleswerefoundinthis
areabefore, mainlybecauseof theabsenceof natural cavesandoftheprevalenceof
250 Shun-lobi UENoand Takao NAITo
graniticgroundnot favourablefor theexistenceof upperhypogean trechines.
In May of this year, thesecondauthorof thepresent paper made two tripsto
southwesternShikokuandsucceededinfindingoutthreenewlocalitiesofYamautidius
at the southeastern side of the Uwa Hills to the west of the middle course of the
Shimanto-gawaRiver. Ateachlocality,anophthalmictrechinesweredugout fromthe
upperhypogean zone,andthegeneric rangewasextendedsouthwestwardsformore
than30kminabeeline,sothatboththelong-pendingproblemsmentionedabovewere
clarifiedbythis discovery. After acloseexamination, it becameapparent that the
specimensobtainedwereclassifiedintotwonewspeciesoftheuozumii complexofthe
1atppennisgroup. Theywillbedescribedinthepresentpaperunder thenamesY.croson
andY.sucmo. Theabbreviationsusedarethesameasthoseexplainedonp7ofUtNo's
1982paper.
Yamautidius(s.str)crosonS.UtNoetNAIT0,sp
nov
(Figs.1-3)
Length: 3.35-3.60mm (fromapicalmarginofclypeustoapicesof elytra).
Related toY. uozumiiand identical withthelatter in thecharacteristicconforma-
tion of thebasal parts of elytra, but a little larger and dark-coloured, with
m ore
anteriorlyenlargedprothoraxandobviouslyamplerelytra. Definitelydifferent fromY.
uozumii inconfigurationofmalegenitalia,particularlyofcopulatorysclerite.
Colourreddishbrown,shiny,withinfuscatedheadandpronotum;clypeus,labrum,
palpi,antennae,venter ofhindbody,andlegspaler thanelytra. Microsculpturefine,
distinctlyimpressedonheadandpartiallydegeneratedonpronotum,mostlyconsisting
oftransversemeshes,largelyobliteratedonelytrathoughperceptiblehereandthereas
finetransverselines.
Headfairlylarge,similar tothatofY.uozumii;antennaerelativelylong, reaching
apicaltwo-fifthsofelytrain ,withpedicelaboutahalfaslongasantennomere3or4,
eachofwhichisnearly3.5timesaslongaswide,antennomeres8-10eachsubcylindrica1
and nearly three times as long as wide, terminal antennomere about as long as
antennomere3andobviouslylongerthanscape.
Pronotumwiderthanheadandalittlewiderthanlong,widestataboutsix-sevenths
from base, andalmoststraightlynarrowedposteriadtowardsante-basalsinuationof
sides;PW/HW130-1.38 (M I33),PW/PL1.03-1.10 (M1.06),PW/PA t29-1.34
(M I32),PW/PB l66-1.76(M I71);sidesratherwidelyreflexednear frontangles,
verynarrowlyborderedatmiddle,andmoderatelyreflexedinbasalarea,brieflybut
stronglyroundedin front,eitherstraightorslightlyemarginateatmiddle,shallowly
sinuateataboutbasalsixth,andthenmoreorlessdivergenttowardshindangles,which
areacuteandpostero-1atera11yprotrudent;apexmuchwiderthanbase,nearlystraightor
slightlybisinuate,PA/PBl25-1.35(MI29),withfrontanglesproducedforwardsand
roundedat theexternalsidestogetherwithanteriormost portionsof lateralmargins;
baseslightlyemarginateat middleandslightlysinuateoneachsideinsidehindangle;